As far as the question of who fits the bill in the article, it literally says the players who couldn't do the physical test were placed on the PUP list until they could pass it. PUP list at the time:
Lucas, Baker and Dodson who were all injured
Boykn, Reed II, Drake Thomas, Cam Young, Bradford, Gibbs, DJ James, Pritchett and Dee Williams.
My guess would be Bradford since he was non-football injury designation, but it could also be Cam Young.
I can't imagine LBs, a safety, CBs or a WR having difficulty doing shuttles.